James McCullen Destro XXIV is the man behind M.A.R.S. Industries, the undisputed leader in providing high-tech weapons to world powers…for the right price.

But the emergence of Energon has changed everything.

As Destro’s ambitions grow, the "Crimson Twins" Tomax and Xamot Paoli emerge to destroy their competition, and Cobra Commander realizes his current ally could be his future greatest enemy.